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@media(prefers-reduced-motion:reduce){html{scroll-behavior:auto}}
@media print{body{background:white;color:black;font-size:11pt}.topline,.nav,.menu-button,.toc,.footer,.related,.skip{display:none}.wrap{max-width:none;padding:0}.masthead{padding:0 0 15px}.seal{width:40px}.brand-name{font-size:15pt}.article-head h1{font-size:28pt}.article-layout{display:block}.article-body{font-size:11pt;line-height:1.5}.article-body h2{font-size:18pt}figure,table,.abstract{break-inside:avoid}a{color:black}.article-head{padding-top:15px}.article-body figure img{max-height:300px;object-fit:contain}.breadcrumb{display:none}}

/* Contributor identities and featured research */
.author-byline{display:flex;align-items:center;gap:17px;margin:27px 0 15px}.author-portrait{width:74px;height:88px;object-fit:cover;flex:none;border:1px solid var(--line)}.author-byline .eyebrow{display:block;font-size:13px;letter-spacing:1px}.author-byline .eyebrow a{font-size:17px;letter-spacing:0;text-transform:none}.author-title,.author-affiliation{display:block;font:13px/1.6 Arial;color:var(--muted)}.author-initials{display:flex;align-items:center;justify-content:center;background:#e0e6e1;font:26px Georgia;color:var(--ink)}.paper-card .card-byline{font:12px/1.6 Arial;margin:0 0 17px}.archive-byline{font:13px/1.6 Arial!important;margin-top:12px!important}.author-bio{border-top:1px solid var(--line);padding-top:15px;margin-top:35px;font:15px/1.75 Arial}.author-bio h2{font-size:25px}.featured-research{display:grid;grid-template-columns:1fr 1fr;gap:45px;border-bottom:1px solid var(--line);align-items:center}.featured-research h2{font-size:39px}.featured-research h2 a{color:var(--ink);text-decoration:none}.featured-text>p{color:var(--muted);font-size:17px}.feature-figure{border:1px solid var(--line)}.feature-figure img{width:100%}.feature-author{display:flex;align-items:center;gap:16px;margin:22px 0}.feature-author a{font-weight:bold;font-size:16px}.feature-author span{display:block;font:12px/1.6 Arial;color:var(--muted)}.profile-layout{display:grid;grid-template-columns:260px minmax(0,780px);gap:60px;padding-bottom:45px}.profile-portrait{width:260px;max-width:100%;height:auto;border:1px solid var(--line)}.author-initials.profile-portrait{height:280px;font:65px Georgia}.profile-image{padding-top:20px}.profile-layout h2:first-child{margin-top:14px}
@media(max-width:700px){.featured-research,.profile-layout{grid-template-columns:1fr;gap:25px}.featured-research h2{font-size:32px}.profile-image{max-width:230px}.author-byline{align-items:flex-start}.author-affiliation{max-width:270px}.profile-portrait{max-height:350px;object-fit:cover}.feature-figure{display:none}}
@media print{.featured-research{display:block}.author-portrait{width:45px;height:55px}.author-byline{margin:10px 0}.author-bio{break-inside:avoid}.profile-layout{display:block}.profile-image{max-width:150px}}

.author-byline img.author-portrait{object-position:center 28%}

.taxonomy-summary{padding:22px 26px;background:#eeeee5;border-left:4px solid var(--gold);margin:26px 0;font:15px/1.7 Arial}.taxonomy-summary strong{font-size:20px}.article-body .taxonomy-rule{font:14px/1.8 Arial;color:var(--muted);border-left:2px solid var(--line);padding-left:17px}.article-body li{margin-bottom:9px}

/* EG-26-002 specimen plates and resource maps */
.pumpkin-paper{min-width:0}.pumpkin-paper .spice-plate{margin:32px 0;background:#f5f1e7;border:1px solid var(--line)}.pumpkin-paper .spice-plate>a{display:block}.pumpkin-paper .spice-plate img{display:block;width:100%;height:auto;max-height:none;object-fit:contain}.pumpkin-paper .spice-plate figcaption{padding:15px 18px;margin:0;font:13px/1.65 Arial,sans-serif;color:var(--muted)}.plate-link{display:inline-block;margin-left:5px;white-space:nowrap}.pumpkin-paper .table-scroll{max-width:100%;overflow-x:auto}.pumpkin-paper table{min-width:610px}.pumpkin-paper td,.pumpkin-paper th{vertical-align:top}.pumpkin-paper strong{color:var(--ink)}
@media(max-width:600px){.pumpkin-paper .spice-plate{margin-left:-8px;margin-right:-8px}.pumpkin-paper .spice-plate figcaption{padding:12px}.pumpkin-paper .plate-link{display:block;margin:7px 0 0}}
@media print{.pumpkin-paper .spice-plate img{max-height:360px}.plate-link{display:none}.pumpkin-paper .table-scroll{overflow:visible}.pumpkin-paper table{min-width:0}}

/* Source exhibit: two unaltered screenshot regions, with the complete record beneath. */
.pumpkin-paper .google-exhibit{background:white}.exhibit-head{padding:17px 18px;background:#102838;color:#e6bc78;font:700 12px/1.6 Arial,sans-serif;letter-spacing:1px}.evidence-window{position:relative;overflow:hidden;background:white;margin:15px 10px}.pumpkin-paper .evidence-window img{position:absolute;max-width:none;height:auto;display:block}.evidence-core{aspect-ratio:760/120}.pumpkin-paper .evidence-core img{width:137.7632%;left:-2.6316%;top:-58.3333%}.evidence-optional{aspect-ratio:960/95}.pumpkin-paper .evidence-optional img{width:109.0625%;left:-2.0833%;top:-547.3684%}.evidence-full{margin:16px 0 28px;border:1px solid var(--line);padding:12px 16px;font:14px/1.65 Arial,sans-serif}.evidence-full summary{cursor:pointer;color:var(--ink);font-weight:bold}.evidence-full img{display:block;width:100%;height:auto;margin-top:15px}.evidence-full:not([open]) img{display:none}@media print{.evidence-full{display:none}.pumpkin-paper .evidence-window img{max-height:none}.google-exhibit{break-inside:avoid}}

/* Native-emulsion plate layout */
/* EG-26-002 specimen plates and resource maps */
.ranch-paper{min-width:0}.ranch-paper .spice-plate{margin:32px 0;background:#f5f1e7;border:1px solid var(--line)}.ranch-paper .spice-plate>a{display:block}.ranch-paper .spice-plate img{display:block;width:100%;height:auto;max-height:none;object-fit:contain}.ranch-paper .spice-plate figcaption{padding:15px 18px;margin:0;font:13px/1.65 Arial,sans-serif;color:var(--muted)}.plate-link{display:inline-block;margin-left:5px;white-space:nowrap}.ranch-paper .table-scroll{max-width:100%;overflow-x:auto}.ranch-paper table{min-width:610px}.ranch-paper td,.ranch-paper th{vertical-align:top}.ranch-paper strong{color:var(--ink)}
@media(max-width:600px){.ranch-paper .spice-plate{margin-left:-8px;margin-right:-8px}.ranch-paper .spice-plate figcaption{padding:12px}.ranch-paper .plate-link{display:block;margin:7px 0 0}}
@media print{.ranch-paper .spice-plate img{max-height:360px}.plate-link{display:none}.ranch-paper .table-scroll{overflow:visible}.ranch-paper table{min-width:0}}

